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Equine Locomotion - E-Book

Equine Locomotion - E-Book

Willem Back | Hilary M. Clayton

(2013)

Additional Information

Abstract

The first edition of Equine Locomotion has established itself as the book in the equine literature that discusses all aspects of equine locomotion and gait analysis, written by an international team of editors and contributors.
The new edition continues this trend and gives the reader a complete picture of the horse in motion, at the same time including many recent findings in this area. The book begins with a history of man’s association with the horse and then continues to discuss with comprehensive descriptions of the present state of knowledge beginning with the initiation of gait and ending with the more scientific area of computer modeling. In the new edition, the list of contributors continues to comprise of authors who are acknowledged experts in their subject areas and includes many new illustrations.
• international team of editors and contributors, with leading experts from the USA, the Netherlands, Sweden and France (all centres of excellence for the study of equine locomotion)
• editors are from two of the worlds leading locomotion centres – Utrecht and Michigan
• highly illustrated with nearly 500 detailed line drawings and illustrations
• covers all you will ever need to know about equine locomotion, gait analysis and much more
• international team of editors and contributors, with leading experts from the USA, the Netherlands, Sweden and France (all centres of excellence for the study of equine locomotion)
• editors are from two of the worlds leading locomotion centres – Utrecht and Michigan
• highly illustrated with nearly 500 detailed line drawings and illustrations
• covers all you will ever need to know about equine locomotion, gait analysis and much more

Table of Contents

Section Title Page Action Price
Front cover cover
Equine Locomotion i
Copyright page iv
Table of Contents v
Foreword vii
References viii
Foreword to the first edition ix
References ix
Preface xi
Preface to the first edition xiii
Acknowledgments xv
Contributors xvii
Glossary xxi
1 History 1
Introduction 1
Prehistoric times 1
The ancient cultures 3
The Antique world 3
Through the Dark Ages to the Renaissance 5
From the Renaissance to the 18th century 5
The start of veterinary education 6
The 19th century 6
Muybridge and Marey: revolution in gait analysis 7
German supremacy until World War II 12
The horse in decline 15
The revival in equine locomotion research 17
Equine locomotion research centers and activities 17
Europe 17
Stockholm/Uppsala 17
Vienna 19
Utrecht 19
Alfort 20
Zurich 21
United Kingdom 21
Belgium 22
Other places in Europe 22
North America 22
Saskatoon 22
Michigan 22
Missouri 23
Other places in North America 23
Elsewhere in the world 24
Concluding remarks 24
References 24
2 Measurement techniques for gait analysis 31
Introduction 31
Interpreting the effects of biological variability 31
Treadmill evaluation 32
Kinematic analysis 33
Markers 33
Calibration 34
Sampling frequency 35
Digitization 35
Smoothing 35
Transformation 36
Normalization 36
Kinematic data 37
Sagittal plane analysis 37
Multi-planar analysis 37
Three-dimensional analysis 37
Methods of kinematic analysis 38
Electrogoniometry 38
Photographic systems 39
Optoelectronic systems 39
Electromagnetic systems 40
Ultrasonographic system 41
Kinetic analysis 41
Ground reaction force 41
Normalization 43
Force plate 43
Force shoes 45
Indirect methods of GRF measurement 46
Inverse dynamic analysis 47
Tissue strain 47
Strain transducers 47
Measuring strains in hard tissues 47
Measuring strains in tendons and ligaments 48
Sonomicrometry 49
Calculation of strains 49
Measurement of rein tension 50
Accelerometers, gyroscopes and magnetometers 50
Pressure mats 52
Hoof pressure mats 52
Saddle pressure mats 53
Electromyography 53
EMG equipment 54
The electromyogram 54
New technologies 55
References 55
3 Signals from materials 61
Introduction and definition of signals 61
Choosing the sampling frequency (Nyquist – Shannon sample theorem) 61
Motion of a wheel 61
Resampling and normalization 61
Aliasing – anti-aliasing filter 63
Signal processing 63
Time curve analysis 63
Differentiation 63
Definition of differentiation 63
The physical concept of Newton 63
Phase-plane analysis (practical use of differentiation) 64
Increasing, decreasing and finding a local maximum or the minimum (extreme values) 64
Integration 64
Transformation of the coordinate system 65
Fourier analysis (time domain – frequency domain) 65
Filtering 66
Differentiation in time and frequency domain 67
Integration in time and frequency domain 67
System analysis in time and frequency domain (convolution, special filters) 68
Decision-making 69
Fuzzy logic 69
Artificial neural network 69
References 70
4 Locomotor 73
Introduction 73
Muscles, motors of movements 74
Summary 74
Muscle fiber-types 74
5 Gaits and interlimb coordination 85
Introduction 85
Classification and description of gaits 85
Walk 88
Other walking gaits 88
Trot 88
Pace 91
Canter and gallop 91
Jump 92
Gait transitions 92
Velocity-related changes in stride variables 93
Gait development and training effects 94
Influence of the treadmill on gait characteristics 94
Locomotion and respiratory coupling 95
References 96
6 Forelimb function 99
Terminology 99
Musculotendinous architecture 99
Extrinsic muscles of the forelimb 101
Intrinsic muscles of the forelimb 103
Sagittal plane analysis of forelimb kinematics and kinetics 105
The trot 106
Scapula 106
Shoulder joint 106
Elbow joint 106
Carpal joint 106
Metacarpophalangeal (MCP) joint 108
Distal interphalangeal (DIP) joint 109
Functional interpretation 110
The walk 111
The canter 113
The gallop 114
Three-dimensional kinematics 115
Three-dimensional forelimb kinematics for locomotion at walk and trot in a straight line 116
Distal interphalangeal joint 116
Proximal interphalangeal joint 119
Metacarpophalangeal joint 119
Carpal joint 120
Three-dimensional forelimb kinematics when turning sharply at walk 120
Landing (from first impact to hoof stabilization) 120
Bearing phase (from hoof stabilization to heel-off) 120
Breakover (from heel-off to toe-off) 122
References 122
7 Hind limb function 127
Introduction 127
Musculotendinous architecture 127
The stay apparatus 130
Sagittal plane analysis of hind limb kinematics and kinetics 131
The trot 133
Hip joint 133
Stifle joint 135
Tarsal joint 135
Metatarsophalangeal (MTP) joint 136
Distal interphalangeal (DIP) joint 137
Hoof interaction with the ground 137
The walk 138
The canter 140
Three-dimensional kinematics of the tarsal joint at trot 142
References 143
8 The role of the hoof and shoeing 147
Introduction 147
Functional anatomy of the foot 147
General anatomy 147
Vascular supply 148
Nerve supply 148
Proprioception 148
Hoof mechanics in the standing horse 148
Hoof–pastern axis 148
Hoof balance 148
Craniocaudal balance 148
Mediolateral balance 150
Four point trimming 152
Hoof mechanics during locomotion 152
Initial ground contact 152
Impact 152
Hoof mechanism 156
Center of pressure 156
Breakover 157
Flight arc 157
Effects of hoof manipulations 158
Hoof angle 158
Hoof length 161
Effects of shoe manipulations 161
Shoe weight 161
Shoe length 162
Shoe width 163
Hoof pads 163
Toe of the shoe 163
Wedges 164
Side wedges 164
Heel/toe wedges 164
Toe grabs and heel caulks 164
Rims, clips and nails 164
Effect of footing 164
Properties of the ground 164
Practical applications to performance 166
Interference 166
Action 166
Practical application to lameness 166
Hoof cracks 166
P3 fracture 166
Laminitis 166
Navicular syndrome 167
Arthrosis and arthritis 167
Spavin 168
Patellar fixation 168
Flexural limb deformities 168
Angular limb deformities 168
Conclusion 169
References 169
9 Gait adaptation in lameness 175
Introduction 175
Definitions 175
Classification of lameness 175
Kinematics of lameness 176
Supporting limb lameness 176
Temporal stride pattern 176
Summary 177
Hoof movement 177
Summary 178
Limb movement and joint angle patterns 178
Summary 179
Head and trunk movement 180
Compensatory lameness 181
Summary 182
Swinging limb and mixed lameness 182
10 The neck and back 199
Function 199
Anatomy 199
Bones 199
Vertebral joints 201
Intervertebral discs 201
Ligaments 201
Muscles and nerves 201
Biomechanics 205
Coordination and locomotion 206
Walk 206
Trot 207
Neck muscle activity and neck orientation 208
Effect of neck orientation on neck muscle activity 209
Trunk muscle activity and thoracolumbar movements 209
Canter 209
Walk vs trot vs canter 211
Neck dysfunction 211
Effect of different head and neck positions on physiology and performance during exercise 212
Discussion of measurements from the neck 216
Back dysfunction 216
Introduction 216
Kinematic examination of normal horses 216
Kinematic examination of horses with back pain 218
Differences between horses with and without back pain 218
The effect of head and neck position on the movement of the back 219
The effect of local analgesia in the movement of the back 220
General conclusions 220
References 224
11 The effects of conformation 229
Introduction 229
The study of equine conformation 229
History and tradition 229
Subjective evaluation 229
Quantitative analysis 230
Conformation and growth 230
Conformation, performance and soundness 232
Deviations of limb and toe axis 232
Head, neck and body 234
Forelimbs 236
Hind limbs 238
Predicting performance and soundness by conformation analysis 241
References 242
12 Genetic contributions to exercise and athletic performance 245
Introduction 245
Historical background and the pre-Mendelian era 245
Selection for locomotor performance 246
New genomics technologies for trait mapping and gene expression 246
Selection for locomotor soundness 251
Conformational lameness-causing disorders 251
Neuromuscular lameness-causing disorders 253
Degenerative lameness-causing disorders 254
Developmental lameness-causing disorders 254
Complex chondrodysplasia: osteochondrosis 254
Simple chondrodysplasia: dwarfism 257
Conclusions 259
References 259
13 The response of musculoskeletal tissues to exercise 267
General introduction 267
Articular cartilage 268
Introduction 268
Basic principles of articular cartilage biology 268
Developmental aspects of articular cartilage 270
The effect of exercise on articular cartilage during growth and development 271
The effect of exercise on articular cartilage in young adult animals 273
Conclusion 274
Bone 274
Introduction 274
Morphology and physiology of bones 274
General response of bone to exercise 275
The need for exercise 275
The response of bone to exercise in the horse 276
Conclusion 278
Muscles 278
Overview 278
A synopsis of equine muscle exercise physiology 279
Muscular adaptations to training 284
Muscle fiber size 284
Muscle fiber type transitions 284
Metabolic changes and increased capillary density 284
Physiological adaptations and buffering capacity 286
Other training consequences 286
Overtraining 286
Detraining 286
Possible mechanisms underlying muscular adaptations to training 287
Implications of training-induced changes to the physiologic response to exercise 288
Tendons and ligaments 290
Introduction 290
Tendon composition and structure 291
Tendon biomechanics 292
Effects of exercise 293
The effects of ageing on tendon tissue 294
The effects of exercise on tendon tissue 294
Conclusion 295
General conclusion 296
References 296
14 Performance in equestrian sports 305
Introduction 305
Racing sports 305
Speed 305
Stride length 305
Stride frequency 305
Efficiency of movement 306
Quarter Horse racing 306
Stride variables 306
Sidedness 307
Effect of urging 307
Thoroughbred racing 307
Stride variables 307
Head and neck motion 308
Lead changes 308
Acceleration 308
Ground reaction forces 308
Limb kinetics 308
Fatigue 309
Injury 309
Training 309
Track surface 309
Standardbred racing 309
Trotters 309
Stride variables 309
Reproducibility of gait 310
Ground reaction forces 310
Sidedness 310
Effect of track design 311
Conformation of trotters 311
Pacers 311
Stride variables 311
Sidedness 311
Conformation 311
Endurance racing 311
Sport horses 312
Quality of movement 312
Stride and swing duration 312
Forelimb kinematics 312
Hind limb kinematics 312
Breed differences 313
Ontogeny of gait 314
Normalization of temporal variables 315
Linear similarity 315
Dynamic similarity 315
Coordination 315
Effect of training 316
Effect of a rider 320
Dressage 320
Rider effects 321
The gaits 322
The walk 322
Half pirouette at the walk 323
The trot 323
The passage 324
The piaffe 324
The canter 325
Lead changes at the canter 326
The canter pirouette 326
The capriole 327
Transitions 327
Conformation of dressage horses 327
Jumping 327
Terminology 328
Jumping mechanics 328
The approach and lift-off 328
The jump suspension 330
The landing and departure 331
Water jump technique 332
Puissance jumping technique 332
Conformation of jumping horses 332
The effect of early training on jumping ability and the predictability of jumping performance 332
The effect of a rider 333
Eventing 333
Dressage performance 333
Speed and endurance performance 333
Effect of added weight on jumping performance 334
Western sports 334
Stability and maneuverability 334
Reaction time 334
Western pleasure gaits 335
Cutting 335
References 336
15 Horse–rider interaction 341
Introduction 341
Saddle 341
The electronic saddle mat 341
Saddle pressure in the standing horse 341
The effect of the saddle on motion of the back 342
Saddle types 343
Saddle fit and saddle pads 343
Panels and girths 343
Bits and reins 343
Bits 343
Reins 345
Rein tension 345
Basic rider movements 347
Trot 347
Rider movements at sitting trot 347
The seat at rising trot 348
Walk 349
Rider movements in the sagittal plane at walk 349
Lateral and rotational rider movements at walk 350
Canter 353
Rider skill 353
General 353
Trot 353
Walk 354
The hand 355
Horse movement in relation to rider skill 355
The influence of the rider’s weight 357
Load carrying at walk 357
Load carrying at trot 357
Rider weight 357
Riding methodology 357
Rhythm 357
Suppleness/relaxation 358
Contact 358
Impulsion 359
Straightness 359
Asymmetry basics 359
Asymmetry in rising trot 359
Collection 360
General 360
Head and neck positions 361
Walk 361
Trot 361
Forward stepping of the hind limbs relative to the hind quarters 364
Hind limb protraction–retraction? 364
Over-track distance 364
Lowering of the hind quarters, and increased hind joint angulation 365
Diagonal dissociation 365
Relative weight bearing 365
Absolute versus relative weight 365
References 366
16 Ethology and welfare aspects 369
Introduction 369
Development of locomotion 369
Early kinetic activity 369
Grazing 370
Play 371
Development of laterality (sidedness) 371
Natural rhythms of activity 372
The impact of intensive management on locomotion 372
Feeding 372
Impact of housing/restricted grazing on locomotion 373
Effect of management regimes on locomotion and welfare 373
Impact of exercise on welfare 374
Overtraining 374
Problems with locomotory behavior during training or performance 375
Locomotion and the impact of different training methods 375
Head and neck positions 375
Breeding or training certain types of locomotory responses and welfare implications 376
Concluding remarks – new approaches including equitation science 377
References 377
17 Rehabilitation of the locomotor apparatus 381
General introduction 381
Historical background: rehabilitation defined 381
Longevity and musculoskeletal disorders 382
Prevalence of neck and back pain 384
Patho-anatomical diagnosis 386
Functional assessment 387
Objective measurements 388
Kinematics 388
EMG 388
Pressure algometry 390
Ultrasonography 394
Rehabilitation techniques: manual therapy 397
Evaluation of information on manipulative/mobilization intervention 398
Evaluation of information on exercise and facilitation techniques 399
Exercise-based treatment techniques: motor control 399
Facilitation-based exercise therapy 401
Hydrotherapy 404
Evaluation of information on mechanical and electro-physical intervention 405
Acupuncture 407
Conclusions 408
References 409
18 Metabolic energetics 419
Introduction 419
Production of energy 420
Aerobic phosphorylation 420
Aerobic glycolysis 420
Fatty acid utilization 422
Anaerobic phosphorylation 422
Phosphocreatine reaction 422
Myokinase reaction 422
Anaerobic glycolysis 422
Regulation of aerobic and anaerobic pathways 422
Energy pathway contributions in the exercising horse 423
Energy substrates 423
Carbohydrates 424
Fat 424
Protein 425
Effects of dietary alterations on energy substrate utilization 425
Energy expenditure 426
Aerobic power 426
Oxygen uptake 426
Oxygen uptake at rest and during submaximal exercise 426
Maximum aerobic power 427
Anaerobic power 427
Anaerobic threshold 429
Postexercise oxygen consumption 429
Economy of locomotion 429
Fatigue 430
Training programs 431
Measuring energy expenditure 431
Treadmills 431
Oxygen consumption 432
Lactate analysis 432
Heart rate 432
Blood gases 433
Blood volume 433
Muscle biopsy 433
Hormone profile 434
Effects of exogenous GH 434
Endogenous GH responses to exercise 435
Thermoregulatory consequences of exercise 435
Conclusions 436
References 436
19 Mechanical analysis and scaling 443
Mechanical analysis 443
Introduction 443
Inverse dynamic analysis 444
Linked segment model 444
Inverse dynamic calculations 444
Measurement of input variables 446
Net joint moment and muscle force 447
Net joint force and joint contact force 450
Continued calculations: power 450
Accuracy of inverse dynamic calculations 452
Three-dimensional analysis 453
Formulas for inverse dynamic calculations 454
Hoof segment 454
Pastern segment 455
Velocity and accelerations 455
Symbols 455
Scaled energetics of locomotion 455
Introduction 455
Geometric scaling 455
Differences in proportion 457
Energy cost of locomotion 459
Muscle work and power 460
Maximum aerobic capacity 462
Concluding remarks 463
References 463
20 Modeling, simulation and animation 467
Development of a computer model 467
Introduction 467
History of computer modeling 467
Computer modeling and biomechanics 467
Applications of computer models 467
Visualization 467
Kinematics (movement) 468
Kinetics (dynamics) 469
Finite element analysis 470
Development of a computer model: a horse history 472
Development of a computer simulation: a human perspective 474
Introduction 474
Designing musculoskeletal models 474
Practical applications 476
Development of computer animation: an equine perspective 476
Introduction 476
History 477
Modern animation 478
Motion capture 478
Optical systems 479
Non-optical systems 479
References 485
Index 489
A 489
B 489
C 490
D 491
E 491
F 492
G 493
H 493
I 494
J 495
K 495
L 495
M 496
N 497
O 497
P 498
Q 498
R 498
S 499
T 500
U 501
V 501
W 502
X 502
Z 502